Family of beetles
Endomychidae , or handsome fungus beetles , is a family of beetles with representatives found in all biogeographic realms . There are around 120 genera and 1300 species. The family was established based on the type genus Endomychus , a genus erected in 1795 by Panzer which was applied to a species that Linnaeus called Chrysomela coccinea . As the common name suggests, Endomychidae feed on fungi . Crowson , in his influential treatment of the beetles, placed the family within the Cucujoidea . They have a tarsal formal of 4-4-4 or 3-3-3 and the wings lack a closed radial cell. The second antennal segment has a sensory appendage that is as long as the third antennal segment. The family has also been grouped with the Coccinellidae in a group called the Trimera for having pseudotrimerous tarsi.[ 1] A 2015 molecular phylogeny study found that the Cucujoidea were found to be non-monophyletic and the Endomychidae was refined with the removal of the Anamorphinae from within the family and elevated to the status of a full family, Anamorphidae . Mycetaeinae and Eupsilobiinae were also found not to belong within the clades of the core Endomychidae, and likewise reclassified into the families Mycetaeidae and Eupsilobiidae .[ 2]
The subfamilies that are included:
Genera
These 70 genera belong to the family Endomychidae as defined in the past (due to revisions, this may not be entirely accurate):
Acinaces Gerstaecker, 1858 g
Aclemmysa Reitter, 1904 g
Agaricophilus Motschulsky, 1838 g
Amphisternus Germar, 1843 g
Amphistethus Strohecker, 1964 g
Ancylopus Costa, 1854 g
Anidrytus Gerstaecker, 1858 i c g
Aphorista Gorham, 1873 i c g b
Archipines Strohecker, 1953 i c g
Asymbius Gorham, 1896 g
Atopomychus Tomaszewska & Szawaryn, 2013 g
Atrichonota Arrow, 1925 g
Avencymon Strohecker, 1971 g
Beccariola Arrow, 1943 g
Bolbomorphus Gorham, 1888 g
Brachytrycherus Arrow, 1920 g
Bystodes Strohecker, 1953 g
Cholovocera Motschulsky, 1838 g
Chondria Gorham, 1888 g
Corynomalus Chevrolat in Dejean, 1836 i c g
Cyclotoma Mulsant, 1851 i g
Danae Reiche, 1847 i c g b
Danascelis Tomaszewska, 1999 i c g
Dapsa Latreille, 1829 g
Dexialia Sasaji, 1970 g
Dialexia Gorham, 1891 g
Discolomopsis Shockley, 2006 g
Displotera Reitter, 1877 g
Ectomychus Gorham, 1888 g
Eidorus g
Endomychus Panzer, 1795 i c g b
Ephebus Chevrolat in Dejean, 1836 i c g
Epipocus Germar, 1843 i c g b
Epopterus Chevrolat in Dejean, 1836 i c g
Eucteanus Gerstaecker, 1857 g
Eucymon g
Eumorphus Weber, 1801 g
Geoendomychus Lea, 1922 g
Glesirhanis Shockley & Alekseev, 2014 g
Hadromychus Bousquet & Leschen, 2002 i c g b
Holoparamecus Curtis, 1833 i c g b
Hylaia Guerin, 1857 g
Hylaperdina Tomaszewska, 2012 g
Idiophyes Blackburn, 1896 g
Indalmus Gerstaecker, 1858 g
Leiestes Chevrolat, 1836 g
Lycoperdina Latreille, 1807 i c g b
Meilichius Gerstaecker, 1857 g
Merophysia Lucas, 1852 g
Microxenus Wallaston, 1861 g
Mycetina Mulsant, 1846 i c g b
Mychothenus Strohecker, 1953 g
Natalinus Tomaszewska, 2011 g
Ohtaius Chûjô, 1938 g
Palaeoestes Kirejtshuk & Nel, 2009 g
Paniegena Heller, 1916 g
Parasymbius Arrow, 1920 g
Phymaphora Newman, 1838 i c g b
Phymaphoroides Motschoulsky, 1857 g
Pleganophorus Hampe, 1855 g
Polymus Mulsant, 1846 g
Pseudindalmus Arrow, 1920 g
Rhanidea Strohecker, 1953 i c g b
Saula Gerstaecker, 1858 i c g
Sinocymbachus Strohecker & Chujo, 1970 g
Stenotarsus Perty, 1832 i c g b
Stethorhanis Blaisdell, 1931 i c g
Trochoideus Westwood, 1833 i c g b
Trycherus Gerstaecker, 1857 g
Xenomycetes Horn, 1880 i c g b
Data sources: i = ITIS,[ 3] c = Catalogue of Life,[ 4] g = GBIF,[ 5] b = Bugguide.net[ 6]
Fossil genera
Burmalestes Tomaszewska and Ślipiński, 2018[ 7] Burmese amber , Myanmar, Late Cretaceous (Cenomanian )
Cretolestes Tomaszewska, Ślipiński and Ren, 2018[ 7] Burmese amber, Myanmar, Cenomanian
Cretaparamecus Tomaszewska, Ślipiński, Bai and Zhang, 2018[ 7] Burmese amber, Myanmar, Cenomanian
Palaeomycetes Tomaszewska, Ślipiński and Ren, 2018[ 7] Burmese amber, Myanmar, Cenomanian
Zemyna Tomaszewska, 2018[ 7] (replaced name Laima Alekseev and Tomaszewska, 2018) : Baltic amber , Eocene
References
^ Tomaszewska, K.Wioletta (2000). "Morphology, phylogeny and classification of adult endomychidae (Coleoptera: Cucujoidea)" (PDF) . Annales Zoologici . 50 (4): 449– 558.
^ Robertson, James A; Ślipiński, Adam; Moulton, Matthew; Shockley, Floyd W; Giorgi, Adriano; Lord, Nathan P; McKenna, Duane D; Tomaszewska, Wioletta; Forrester, Juanita; Miller, Kelly B; Whiting, Michael F; McHugh, Joseph V (2015). "Phylogeny and classification of Cucujoidea and the recognition of a new superfamily Coccinelloidea (Coleoptera: Cucujiformia)" (PDF) . Systematic Entomology . 40 (4): 745. doi :10.1111/syen.12138 . S2CID 55206626 .
^
"Endomychidae Report" . Integrated Taxonomic Information System . Retrieved 2018-05-07 .
^ "Browse Endomychidae" . Catalogue of Life . Archived from the original on 2021-06-18. Retrieved 2018-05-07 .
^
"Endomychidae" . GBIF . Retrieved 2018-05-07 .
^
"Endomychidae Family Information" . BugGuide.net . Retrieved 2018-05-07 .
^ a b c d e Tomaszewska, Wioletta; Ślipiński, Adam; Bai, Ming; Zhang, Weiwei; Ren, Dong (November 2018). "The oldest representatives of Endomychidae (Coleoptera: Coccinelloidea) from the Upper Cretaceous Burmese amber" . Cretaceous Research . 91 : 287– 298. doi :10.1016/j.cretres.2018.07.001 . S2CID 135401188 .
External links