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Systemic risk

In finance, systemic risk is the risk of collapse of an entire financial system or entire market, as opposed to the risk associated with any one individual entity, group or component of a system, that can be contained therein without harming the entire system.[1][2][3] It can be defined as "financial system instability, potentially catastrophic, caused or exacerbated by idiosyncratic events or conditions in financial intermediaries".[4] It refers to the risks imposed by interlinkages and interdependencies in a system or market, where the failure of a single entity or cluster of entities can cause a cascading failure, which could potentially bankrupt or bring down the entire system or market.[5] It is also sometimes erroneously referred to as "systematic risk".

Explanation

Systemic risk has been associated with a bank run which has a cascading effect on other banks which are owed money by the first bank in trouble, causing a cascading failure. As depositors sense the ripple effects of default, and liquidity concerns cascade through money markets, a panic can spread through a market, with a sudden flight to quality, creating many sellers but few buyers for illiquid assets. These interlinkages and the potential "clustering" of bank runs are the issues which policy makers consider when addressing the issue of protecting a system against systemic risk.[1][6] Governments and market monitoring institutions (such as the U.S. Securities and Exchange Commission (SEC), and central banks) often try to put policies and rules in place with the justification of safeguarding the interests of the market as a whole, claiming that the trading participants in financial markets are entangled in a web of dependencies arising from their interlinkage. In simple English, this means that some companies are viewed as too big and too interconnected to fail. Policy makers frequently claim that they are concerned about protecting the resiliency of the system, rather than any one individual in that system.[6] Systemic risk arises because of the interaction of market participants, and therefore can be seen as a form of endogenous risk.[7]

The risk management literature offers an alternative perspective to notions from economics and finance by distinguishing between the nature of systemic failure, its causes and effects, and the risk of its occurrence.[3] It takes an "operational behaviour" approach to defining systemic risk of failure as: "A measure of the overall probability at a current time of the system entering an operational state of systemic failure by a specified time in the future, in which the supply of financial services no longer satisfies demand according to regulatory criteria, qualified by a measure of uncertainty about the system's future behaviour, in the absence of new mitigation efforts." This definition lends itself to practical risk mitigation applications, as demonstrated in recent research by a simulation of the collapse of the Icelandic financial system in circa 2008.

Systemic risk should not be confused with market or price risk as the latter is specific to the item being bought or sold and the effects of market risk are isolated to the entities dealing in that specific item. This kind of risk can be mitigated by hedging an investment by entering into a mirror trade.

Insurance is often easy to obtain against "systemic risks" because a party issuing that insurance can pocket the premiums, issue dividends to shareholders, enter insolvency proceedings if a catastrophic event ever takes place, and hide behind limited liability. Such insurance, however, is not effective for the insured entity.

One argument that was used by financial institutions to obtain special advantages in bankruptcy for derivative contracts was a claim that the market is both critical and fragile.[1][6][8][9]

Systemic risk can also be defined as the likelihood and degree of negative consequences to the larger body. With respect to federal financial regulation, the systemic risk of a financial institution is the likelihood and the degree that the institution's activities will negatively affect the larger economy such that unusual and extreme federal intervention would be required to ameliorate the effects.[10]

A general definition of systemic risk which is not limited by its mathematical approaches, model assumptions or focus on one institution, and which is also the first operationalizable definition of systemic risk encompassing the systemic character of financial, political, environmental, and many other risks, was put forth in 2010.[11]

The Systemic Risk Centre at the London School of Economics is focused on the study of systemic risk. It finds that systemic risk is a form of endogenous risk, hence frustrating empirical measurements of systemic risk.

Measurement

TBTF/TCTF

According to the Property Casualty Insurers Association of America, there are two key assessments for measuring systemic risk, the "too big to fail" (TBTF) and the "too (inter)connected to fail" (TCTF or TICTF) tests. First, the TBTF test is the traditional analysis for assessing the risk of required government intervention. TBTF can be measured in terms of an institution's size relative to the national and international marketplace, market share concentration, and competitive barriers to entry or how easily a product can be substituted. Second, the TCTF test is a measure of the likelihood and amount of medium-term net negative impact to the larger economy of an institution's failure to be able to conduct its ongoing business. The impact is measure beyond the institution's products and activities to include the economic multiplier of all other commercial activities dependent specifically on that institution. The impact is also dependent on how correlated an institution's business is with other systemic risks.[12]

Too big to fail

The traditional analysis for assessing the risk of required government intervention is the "too big to fail" test (TBTF). TBTF can be measured in terms of an institution's size relative to the national and international marketplace, market share concentration (using the Herfindahl-Hirschman Index for example), and competitive barriers to entry or how easily a product can be substituted. While there are large companies in most financial marketplace segments, the national insurance marketplace is spread among thousands of companies, and the barriers to entry in a business where capital is the primary input are relatively minor. The policies of one homeowners insurer can be relatively easily substituted for another or picked up by a state residual market provider, with limits on the underwriting fluidity primarily stemming from state-by-state regulatory impediments, such as limits on pricing and capital mobility. During the recent financial crisis, the collapse of the American International Group (AIG) posed a significant systemic risk to the financial system. There are arguably either no or extremely few insurers that are TBTF in the U.S. marketplace.

Too connected to fail

A more useful systemic risk measure than a traditional TBTF test is a "too connected to fail" (TCTF) assessment. An intuitive TCTF analysis has been at the heart of most recent federal financial emergency relief decisions. TCTF is a measure of the likelihood and amount of medium-term net negative impact to the larger economy of an institution's failure to be able to conduct its ongoing business.

Network models have been proposed as a method for quantifying the impact of interconnectedness on systemic risk.[13][14][15]

The impact is measured not just on the institution's products and activities, but also the economic multiplier of all other commercial activities dependent specifically on that institution. It is also dependent on how correlated an institution's business is with other systemic risk.[16]

Criticisms of systemic risk measurements

Criticisms of systemic risk measurements: Danielsson et al.[17][18] express concerns about systemic risk measurements, such as SRISK and CoVaR, because they are based on market outcomes that happen multiple times a year, so that the probability of systemic risk as measured does not correspond to the actual systemic risk in the financial system. Systemic financial crises happen once every 43 years for a typical OECD country and measurements of systemic risk should target that probability.

SRISK

A financial institution represents a systemic risk if it becomes undercapitalized when the financial system as a whole is undercapitalized. In a single risk factor model, Brownlees and Engle [19] build a systemic risk measure named SRISK. SRISK can be interpreted as the amount of capital that needs to be injected into a financial firm as to restore a certain form of minimal capital requirement. SRISK has several nice properties: SRISK is expressed in monetary terms and is, therefore, easy to interpret. SRISK can be easily aggregated across firms to provide industry and even country specific aggregates. Last, the computation of SRISK involves variables which may be viewed on their own as risk measures. These are the size of the financial firm, the leverage (ratio of assets to market capitalization), and a measure of how the return of the firm evolves with the market (some sort of time varying conditional beta but with emphasis on the tail of the distribution).

Whereas the initial Brownlees and Engle model is tailored to the US market, the extension by Engle, Jondeau, and Rockinger[20] is more suitable for the European markets. One factor captures worldwide variations of financial markets, another one the variations of European markets. This extension allows for a country-specific factor.

By accounting for different factors, one captures the notion that shocks to the US or Asian markets may affect Europe, but also that bad news within Europe (such as the news about a potential default of one of the countries) matters for Europe. Also, there may be country specific news that does not affect Europe or the US, but matters for a given country. Empirically the last factor is less relevant than the worldwide or European factor.

Since SRISK is measured in terms of currency, the industry aggregates may also be related to Gross Domestic Product. As such one obtains a measure of domestic, systemically important banks.

The SRISK Systemic Risk Indicator is computed automatically on a weekly basis and made available to the community. For the US model, SRISK and other statistics may be found under the Volatility Lab of NYU Stern School website and for the European model under the Center of Risk Management (CRML) website of HEC Lausanne.

Pair/vine copulas

A vine copula can be used to model systemic risk across a portfolio of financial assets. One methodology is to apply the Clayton Canonical Vine Copula to model asset pairs in the vine structure framework. As a Clayton copula is used, the greater the degree of asymmetric (i.e., left tail) dependence, the higher the Clayton copula parameter. Therefore, one can sum up all the Clayton Copula parameters, and the higher the sum of these parameters, the greater the impending likelihood of systemic risk. This methodology has been found to detect spikes in the US equities markets in the last four decades capturing the Oil Crisis and Energy Crisis of the 1970s, Black Monday and the Gulf War in the 1980s, the Russian Default/LTCM crisis of the 1990s, and the Technology Bubble and Lehman Default in the 2000s.[21] Manzo and Picca[22] introduce the t-Student Distress Insurance Premium (tDIP), a copula-based method that measures systemic risk as the expected tail loss on a credit portfolio of entities, in order to quantify sovereign as well as financial systemic risk in Europe.

Valuation of assets and derivatives under systemic risk

Inadequacy of classic valuation models

One problem when it comes to the valuation of derivatives, debt, or equity under systemic risk is that financial interconnectedness has to be modelled. One particular problem is posed by closed valuations chains, as exemplified here for four firms A, B, C, and D:

B might hold shares of A, C holds some debt of B, D owns a derivative issued by C, and A owns some debt of D.[23]

For instance, the share price of A could influence all other asset values, including itself.

The Merton (1974) model

Situations as the one explained earlier, which are present in mature financial markets, cannot be modelled within the single-firm Merton model,[24] but also not by its straightforward extensions to multiple firms with potentially correlated assets.[23] To demonstrate this, consider two financial firms, , with limited liability, which both own system-exogenous assets of a value at a maturity , and which both owe a single amount of zero coupon debt , due at time . "System-exogenous" here refers to the assumption, that the business asset is not influenced by the firms in the considered financial system. In the classic single firm Merton model,[24] it now holds at maturity for the equity and for the recovery value of the debt, that

and

Equity and debt recovery value, and , are thus uniquely and immediately determined by the value of the exogenous business assets. Assuming that the are, for instance, defined by a Black-Scholes dynamic (with or without correlations), risk-neutral no-arbitrage pricing of debt and equity is straightforward.

Non-trivial asset value equations

Consider now again two such firms, but assume that firm 1 owns 5% of firm two's equity and 20% of its debt. Similarly, assume that firm 2 owns 3% of firm one's equity and 10% of its debt. The equilibrium price equations, or liquidation value equations,[25] at maturity are now given by

This example demonstrates, that systemic risk in the form of financial interconnectedness can already lead to a non-trivial, non-linear equation system for the asset values if only two firms are involved.

Over- and underestimation of default probabilities

It is known that modelling credit risk while ignoring cross-holdings of debt or equity can lead to an under-, but also an over-estimation of default probabilities.[26] The need for proper structural models of financial interconnectedness in quantitative risk management – be it in research or practice – is therefore obvious.

Structural models under financial interconnectedness

The first authors to consider structural models for financial systems where each firm could own the debt of other firms were Eisenberg and Noe in 2001.[27] Suzuki (2002) extended the analysis of interconnectedness by modeling the cross ownership of both debt and equity claims.[28] Building on Eisenberg and Noe (2001), Cifuentes, Ferrucci, and Shin (2005) considered the effect of costs of default on network stability.[29] Elsinger's further developed the Eisenberg and Noe (2001) model by incorporating financial claims of differing priority.[30]

Acemoglu, Ozdaglar, and Tahbaz-Salehi, (2015) developed a structural systemic risk model incorporating both distress costs and debt claim with varying priorities and used this model to examine the effects of network interconnectedness on financial stability. They showed that, up to a certain point, interconnectedness enhances financial stability. However, once a critical threshold density of connectedness is exceeded, further increases in the density of the financial network propagate risk.[31]

Glasserman and Young (2015) applied the Eisenberg and Noe (2001) to modelling the effect of shocks to banking networks. They develop general bounds for the effects of network connectivity on default probabilities. In contrast to most of the structural systemic risk literature, their results are quite general and do not require assuming a specific network architecture or specific shock distributions.[32]

Risk-neutral valuation: price indeterminacy and open problems

Generally speaking, risk-neutral pricing in structural models of financial interconnectedness requires unique equilibrium prices at maturity in dependence of the exogenous asset price vector, which can be random. While financially interconnected systems with debt and equity cross-ownership without derivatives are fairly well understood in the sense that relatively weak conditions on the ownership structures in the form of ownership matrices are required to warrant uniquely determined price equilibria,[23][28][30] the Fischer (2014) model needs very strong conditions on derivatives – which are defined in dependence on any other liability of the considered financial system – to be able to guarantee uniquely determined prices of all system-endogenous liabilities. Furthermore, it is known that there exist examples with no solutions at all, finitely many solutions (more than one), and infinitely many solutions.[23][25] At present, it is unclear how weak conditions on derivatives can be chosen to still be able to apply risk-neutral pricing in financial networks with systemic risk. It is noteworthy, that the price indeterminacy that evolves from multiple price equilibria is fundamentally different from price indeterminacy that stems from market incompleteness.[25]

Factors

Factors that are found to support systemic risks[33] are:

  1. Economic implications of models are not well understood. Though each individual model may be made accurate, the facts that (1) all models across the board use the same theoretical basis, and (2) the relationship between financial markets and the economy is not known lead to aggravation of systemic risks.
  2. Liquidity risks are not accounted for in pricing models used in trading on the financial markets. Since all models are not geared towards this scenario, all participants in an illiquid market using such models will face systemic risks.

Diversification

Risks can be reduced in four main ways: avoidance, diversification, hedging and insurance by transferring risk. Systematic risk, also called market risk or un-diversifiable risk, is a risk of a security that cannot be reduced through diversification. Participants in the market, like hedge funds, can be the source of an increase in systemic risk[34] and the transfer of risk to them may, paradoxically, increase the exposure to systemic risk.

Until recently, many theoretical models of finance pointed towards the stabilizing effects of a diversified (i.e., dense) financial system. Nevertheless, some recent work has started to challenge this view, investigating conditions under which diversification may have ambiguous effects on systemic risk.[35][36] Within a certain range, financial interconnections serve as a shock-absorber (i.e., connectivity engenders robustness and risk-sharing prevails). But beyond the tipping point, interconnections might serve as a shock-amplifier (i.e., connectivity engenders fragility and risk-spreading prevails).

Regulation

One of the main reasons for regulation in the marketplace is to reduce systemic risk.[6] However, regulation arbitrage – the transfer of commerce from a regulated sector to a less regulated or unregulated sector – brings markets a full circle and restores systemic risk. For example, the banking sector was brought under regulations in order to reduce systemic risks.[37] Since the banks themselves could not give credit where the risk (and therefore returns) were high, it was primarily the insurance sector which took over such deals. Thus the systemic risk migrated from one sector to another and proves that regulation of only one industry cannot be the sole protection against systemic risks.[38]

Project risks

In the fields of project management and cost engineering, systemic risks include those risks that are not unique to a particular project and are not readily manageable by a project team at a given point in time. They are caused by micro or internal factors i.e. uncertainty resulting from attributes of the project system/culture. Some use the term inherent risk. These systemic risks are called individual project risks e.g. in PMI PMBOK(R) Guide. These risks may be driven by the nature of a company's project system (e.g., funding projects before the scope is defined), capabilities, or culture. They may also be driven by the level of technology in a project or the complexity of a project's scope or execution strategy.[39] One recent example of systemic risk is the collapse of Lehman Brothers in 2008, which sent shockwaves throughout the financial system and the economy.[40] In contrast, those risks that are unique to a particular project are called overall project risks aka systematic risks in finance terminology. They are project-specific risks which are sometimes called contingent risks, or risk events. These systematic risks are caused by uncertainty in macro or external factors of the external environment. "The Great Recession" of the late 2000s is an example of systematic risk.[40] Overall project risks are determined using PESTLE, VUCA, etc.

PMI PMBOK(R) Guide defines individual project risk as "an uncertain event or condition that, if it occurs, has a positive or negative effect on one or more project objectives," whereas overall project risk is defined as "the effect of uncertainty on the project as a whole … more than the sum of individual risks within a project, since it includes all sources of project uncertainty … represents the exposure of stakeholders to the implications of variations in project outcome, both positive and negative."[41]

Systemic risk and insurance

In February 2010, international insurance economics think tank, The Geneva Association, published a 110-page analysis of the role of insurers in systemic risk.[42]

In the report, the differing roles of insurers and banks in the global financial system and their impact on the crisis are examined (See also CEA report, "Why Insurers Differ from Banks").[43] A key conclusion of the analysis is that the core activities of insurers and reinsurers do not pose systemic risks due to the specific features of the industry:

  • Insurance is funded by up-front premia, giving insurers strong operating cash-flow without the requirement for wholesale funding;
  • Insurance policies are generally long-term, with controlled outflows, enabling insurers to act as stabilisers to the financial system;
  • During the hard test of the financial crisis, insurers maintained relatively steady capacity, business volumes and prices.

Applying the most commonly cited definition of systemic risk, that of the Financial Stability Board (FSB), to the core activities of insurers and reinsurers, the report concludes that none are systemically relevant for at least one of the following reasons:

  • Their limited size means that there would not be disruptive effects on financial markets;
  • An insurance insolvency develops slowly and can often be absorbed by, for example, capital raising, or, in a worst case, an orderly wind down;
  • The features of the interrelationships of insurance activities mean that contagion risk would be limited.

The report underlines that supervisors and policymakers should focus on activities rather than financial institutions when introducing new regulation and that upcoming insurance regulatory regimes, such as Solvency II in the European Union, already adequately address insurance activities.

However, during the financial crisis, a small number of quasi-banking activities conducted by insurers either caused failure or triggered significant difficulties. The report therefore identifies two activities which, when conducted on a widespread scale without proper risk control frameworks, have the potential for systemic relevance.

  • Derivatives trading on non-insurance balance sheets;
  • Mis-management of short-term funding from commercial paper or securities lending.

The industry has put forward five recommendations to address these particular activities and strengthen financial stability:

  • The implementation of a comprehensive, integrated and principle-based supervision framework for insurance groups, in order to capture, among other things, any non-insurance activities such as excessive derivative activities.
  • Strengthening liquidity risk management, particularly to address potential mis-management issues related to short-term funding.
  • Enhancement of the regulation of financial guarantee insurance, which has a very different business model than traditional insurance.
  • The establishment of macro-prudential monitoring with appropriate insurance representation.
  • The strengthening of industry risk management practices to build on the lessons learned by the industry and the sharing experiences with supervisors on a global scale.

Since the publication of The Geneva Association statement, in June 2010, the International Association of Insurance Supervisors (IAIS) issued its position statement on key financial stability issues. A key conclusion of the statement was that, "The insurance sector is susceptible to systemic risks generated in other parts of the financial sector. For most classes of insurance, however, there is little evidence of insurance either generating or amplifying systemic risk, within the financial system itself or in the real economy."[44]

Other organisations such as the CEA and the Property Casualty Insurers Association of America (PCI)[45] have issued reports on the same subject.

Discussion

Systemic risk evaluates the likelihood and degree of negative consequences to the larger body. The term "systemic risk" is frequently used in recent discussions related to the economic crisis, such as the Subprime mortgage crisis. The systemic risk of a financial institution is the likelihood and the degree that the institution's activities will negatively affect the larger economy such that unusual and extreme federal intervention would be required to ameliorate the effects. The failing of financial firms in 2008 caused systemic risk to the larger economy. Chairman Barney Frank has expressed concerns regarding the vulnerability of highly leveraged financial systems to systemic risk and the US government has debated how to address financial services regulatory reform and systemic risk.[46][47]

A series of empirical studies published between the 1990s and 2000s showed that deregulation and increasingly fierce competition lowers bank's profit margin and encourages the moral hazard to take excessive credit risks to increase profits.[citation needed] On the other hand, the same effect was measured in presence of a banking oligopoly in which banking sector was dominated by a restricted number of market operators encouraged by their market share and contractual power to set higher loan mean rates.[citation needed] An excessive number of market operators was sometimes deliberately introduced with a below market value selling to cause a price war and a wave of bank massive failures, subsequently degenerating in the creation a market cartel: those two phases had been seen as expressions of the same interest to collude at generally lower prices (and then higher), resulting possible because of a lack of regulation ordered to prevent both of them.[citation needed] Banks are the entities most likely to be exposed to valuation risk as a result of their massive holdings of financial instruments classified as Level 2 or 3 of the fair value hierarchy. In Europe, at the end of 2020 the banks under the direct supervision of the European Central Bank (ECB) held fair-valued financial instruments in an amount of €8.7 trillion, of which €6.6 trillion classified as Level 2 or 3. Level 2 and Level 3 instruments respectively amounted to 495% and 23% of the banks' highest-quality capital (so-called Tier 1 Capital).[48] As an implication, even small errors in such financial instruments' valuations may have significant impacts on banks' capital.

In February 2020 the European Systemic Risk Board warned in a report that substantial amounts of financial instruments with complex features and limited liquidity that sit in banks' balance sheets are a source of risk for the stability of the global financial system.[49] In Europe, at the end of 2020 the banks under the direct supervision of the European Central Bank (ECB) held financial instruments subject to fair value accounting in an amount of €8.7 trillion. Of these, €6.6 trillion were classified as Level 2 or 3 in the so-called Fair Value Hierarchy, which means that they are potentially exposed to valuation risk, i.e. to uncertainty about their actual market value. Level 2 and Level 3 instruments respectively amounted to 495% and 23% of the banks' highest-quality capital (so-called Tier 1 Capital).[50] As an implication, even small errors in such financial instruments' valuations may have significant impacts on banks' capital.

See also

Further reading

  • Website dedicated to systemic risk: http://www.systemic-risk-hub.org/
  • Bartram, Söhnke M.; Brown, Gregory W.; Hund, John (December 2007). "Estimating Systemic Risk in the International Financial System" (PDF). Journal of Financial Economics. 86 (3): 835–869. CiteSeerX 10.1.1.1033.2966. doi:10.1016/j.jfineco.2006.10.001. S2CID 6589252. SSRN 938707.
  • Cont, Rama; Moussa, Amal; Santos, Edson B (December 2013). Network structure and systemic risk in banking systems. pp. 327–368. doi:10.1017/CBO9781139151184. ISBN 9781139151184. {{cite book}}: |journal= ignored (help)
  • Fischer, Tom (2014). "No-arbitrage pricing under systemic risk: Accounting for cross-ownership". Mathematical Finance. 24 (1): 97–124. arXiv:1005.0768. doi:10.1111/j.1467-9965.2012.00526.x. S2CID 153225655. (Published online: 19 Jun 2012)
  • Gray, Dale F. and Andreas A. Jobst, 2011, "Modelling Systemic Financial Sector and Sovereign Risk," Sveriges Riksbank Economic Review, No. 2, pp. 68–106.
  • Gray, Dale F. and Andreas A. Jobst, 2009, "Higher Moments and Multivariate Dependence of Implied Volatilities from Equity Options as Measures of Systemic Risk," Global Financial Stability Report, Chapter 3, April (Washington: International Monetary Fund), pp. 128–31.
  • Gray, Dale F. and Andreas A. Jobst, 2011, "Modeling Systemic and Sovereign Risk," in: Berd, Arthur (ed.) Lessons from the Financial Crisis (London: RISK Books), pp. 143–85.
  • Gray, Dale F. and Andreas A. Jobst, 2011, "Systemic Contingent Claims Analysis – A Model Approach to Systemic Risk," in: LaBrosse, John R., Olivares-Caminal, Rodrigo and Dalvinder Singh (eds.) Managing Risk in the Financial System (London: Edward Elgar), pp. 93–110.
  • Gray, Dale F.; Jobst, Andreas A.; Malone, Samuel (2010). "Quantifying Systemic Risk and Reconceptualizing the Role of Finance for Economic Growth". Journal of Investment Management. 8 (2): 90–110.
  • Hansen, Lars Peter. "Challenges in Identifying and Measuring Systemic Risk" (PDF). National Bureau of Economic Research. Retrieved 6 March 2013.
  • Ilin, Thomas; Varga, Liz (2015). "The uncertainty of systemic risk". Risk Management. 17 (4): 240–275. doi:10.1057/rm.2015.15. S2CID 155209532.
  • Jobst, Andreas A., 2012, "Systemic Risk in the Insurance Sector-General Issues and a First Assessment of Large Commercial (Re-)Insurers in Bermuda," Working paper (March 14). SSRN 2022062.
  • Orlando, Giuseppe; Bufalo Michele; Penikas Henry; Zurlo Concetta (2022). Modern Financial Engineering: Counterparty, Credit, Portfolio and Systemic Risks. World Scientific. ISBN 978-981-125-235-8.
  • Williams, Mark T. (March 2010). "Uncontrolled Risk: The Lessons of Lehman Brothers and How Systemic Risk Can Still Bring Down the World Financial System". Mcgraw-Hill.
  • Zeyu Zheng, Boris Podobnik, Ling Feng and Baowen Li, "Changes in Cross-Correlations as an Indicator for Systemic Risk" (Scientific Reports 2: 888 (2012)).

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